CULTURAL PRACTICES
LEARN
HOW PLANTS GROW
In this chapter and most of
the following chapters, the technical knowledge presented is for your benefit
as a worker with plants. Some of you as
salespersons will also be dealing with customers. In most cases, little will be accomplished
in using technical terms; however, you must understand the technical
terminology that other people use to express it.
It is impossible to cover
completely each of the subject matters to be discussed. Additional, easily available reference
material will be cited for your further enlightenment.
To understand how plants
grow, one must first understand their structures and the functions of the
various parts. This involves roots,
stems, leaves, and reproductive parts.
There is a great variation in appearance of these plant parts as found
throughout the plant kingdom; however, their basic functions remain the same.
ROOTS
In a normal, healthy plant
there is a balance between the shoot system and the root system. This balance is disturbed by root loss during
transplanting or through improper cultivation.
The balance can be restored by pruning some of the top (shoot) growth of
the plant. If this is not done, the
plant may be set back or die because the roots are unable to supply the tops
with adequate moisture and nutrients.
Root loss during transplanting is much greater in bare root trees and
shrubs than in container plants.
There are two kinds of root
systems found in plants, the fibrous root system and the tap root system. The fibrous root system consists of a great
many, much-branched roots that are slender and fiber-like; no one root is more prominent than the others. Most of our nursery plants have this type of
root system.
The tap root system consists
of one main root that grows directly downward, from which branch roots
arise. The main root may be a fleshy,
food storing organ (carrot), or a more or less woody root (oak, citrus). Tap roots generally penetrate much deeper in
the soil than fibrous roots, thus tap rooted plants are much more difficult to
transplant.
The functions of the root system are
absorption, anchorage, conduction and storage.
In land plants, large amounts
of water are absorbed through millions of thin-walled root hairs in close
contact with the soil particles. Each
root hair is formed from a single epidermal cell near the growing tip of the
root. Each one lasts only a short time,
and many are killed during transplanting, causing some plants to wilt because
water loss is greater than intake.
Little water absorption occurs through the epidermis (outer cell layer)
of older roots where root hairs are no longer present. Special care must be given during
transplanting, and roots should be exposed for no longer than absolutely
necessary. In the nursery, they should
be heeled-in with moist sawdust or soil until sold or transplanted. The customer should be told about the effects
of drying the roots of a plant.
Plants that have become
root-bound within their containers also need special attention. The roots should be spread out or pruned if
they are found to be spirally coiled around themselves. Such a plant may strangle itself within a few
years.
Plant nutrients (inorganic
fertilizers) must be in a soluble form so they
can easily pass through the
cell walls into the plant and be carried to the leaves for manufacture into
usable food. Any plant receiving
fertilizer should be well watered so that the fertilizer is diluted throughout
the root zone. An excess of fertilizer
can result in a burning or browning of the foliage. This is a result of water shortage in the
foliage due to its movement out of the plant root when surrounding soil
contains too high a concentration of fertilizer. The foliage is actually dried out.
STEMS
The primary functions of the
stem are support, conduction and storage. Since the stem holds the leaves up,
they receive more light, which is important in the manufacture of plant
food. Some plants have a single stem,
often called a trunk, with various methods of branching. Others have many stems arising from the root
crown. The interior, dense hardwood of
older stems is dead and serves only for support.
Conduction is the moving of
water, mineral solutions and plant food. Water and mineral solutions move
upward from the roots to the leaves through inner tissues called xylem. Manufactured food from the leaves moves out
of the leaves through outer stem tissues called phloem. Between the xylem and the phloem is a tissue
called the cambium layer, which forms new xylem and phloem as may be required
by the plant.
STEM CROSS SECTIONS
Not all plants have the same
arrangement of xylem, phloem and cambium. In monocotyledonous (single seed
leaf) plants, the three tissues are formed in bundles scattered in a
discontinuous vascular system throughout the stem. (Example:
corn, grass).
In the dicotyledonous (2 seed
leaves) plants and gymnosperms (many seed leaves), the xylem is formed on the
inside next to the heartwood, surrounded by a continuous layer of cambium, with
the phloem on the outside. (Example:
most trees, shrubs, annual and perennial plants.) The cambium forms new xylem on the inside as
an annual ring, or cycle of growth, and new layers of phloem on the outside of
the cambium layer. The older layers of
phloem form the bark on trees and shrubs.
It is important to prevent
damage to the cambium layer. This could
be caused by wire too tightly tied to stake up a plant, plastic rope used to
ball up a plant's roots, or mechanical damage such as bruises and cuts. The first two conditions can kill a plant if
unchecked, and the third can be an open invitation for insects and plant
diseases. It may also predispose a plant
to winter injury. When a stem is
completely girdled, the phloem cells below the girdle no longer receive food
from the leaves. Therefore, all leafless
parts below the girdle will be starved.
A girdle is a complete circle of dead tissue around the stem, which
interrupts the supply pipeline.
SPECIALIZED STEMS AND ROOTS
Some plants have stem and
root structures modified so that they hold exceptionally large amounts of
stored food and also function as asexual reproductive structures. Some of the growth patterns in modified
stems are quite complex and involve
changes from vegetative growth to reproductive development and back again. These growth cycles are usually controlled by
changes in temperature, although moisture may be the controlling factor for
tropical plants. Bulbs and corms are
examples of modified stems that function in this way. Some of our most popular horticulture plants
are propagated by the use of bulbs, corms, rhizomes, tubers (modified stems)
and tuberous roots. Lilies, tulips, hyacinths,
narcissus, Dutch iris, and many more early spring flowers develop from
bulbs. Crocus, colchicum, and gladiolus
all have corms. German, or tall, bearded iris; lily-of-the-valley; and orchids
send up leaves and flower stalks from rhizomes.
In addition, grass is planted using sprigs, or stolons,
and strawberry plants are obtained from the ends of runners. Potatoes are planted by using pieces of
tubers. All of these are also modified
stems. Interestingly, all but the
strawberry and potato are members of the monocotyledon subgroup of flowering
plants.
The structure of each of
these is slightly different, but all except tuberous roots have nodes,
comparable to those on a typical stem.
Buds form at these nodes, as on a typical stem. A true bulb, such as a daffodil, consists
primarily of fleshy modified leaves containing stored food. The stem, or basal plate, is a very
compressed structure; e.g., the hard part cut off of the bottom of an onion in
preparing it for cooking is the basal plate.
Each scale leaf has arisen from a node on this plate. Usually a new growing point appears in the axil of about every fourth leaf. In some bulbs a growing point can occur at
each axil (point where a leaf joins the stem). New growing points are initiated in the fall
and normally one continues to develop during spring while the old mother bulb
is flowering. After flowering, the old
bulb gradually disintegrates (in most cases) as the new bulb scales enlarge and
develop. In order to have successful
flowering the next year, foliage leaves must be maintained as long as possible
to provide a source of carbohydrates to be stored in the new bulb. A relatively cool growing season following
bloom is ideal but not what we can ordinarily expect in the Southeast. This explains why commercial production of
bulbs and related structures is not feasible in our area. It is also helpful to feed and water bulbs
following flowering to help maintain vegetative growth as long as
possible. Corms, such as gladiolus,
differ from bulbs in that they are solid stem structures with remains of dried
scale leaves surrounding and protecting them.
Nodes can be seen if these dried leaves are removed. Again, the cycle of changes is controlled by
temperature. During the time the mother corm is flowering, a new corm is
developing at its top. By fall there is
only a small shriveled remnant of the mother.
A period of relatively warm
(not hot) weather is necessary to initiate changes from vegetative growth to
reproductive development within the bulb.
However, the bloom will be produced until after a period of exposure to
temperatures around 40 degrees for about 8 to 12 weeks. Following this, bloom stalks will develop in
bulbs like tulip or daffodil when warm temperatures again prevail. As already mentioned, the changes in some
cases are controlled by differences in moisture. This is typical of species that have
developed in tropical areas where seasons
are wet-dry rather than
hot-cold. The large so-called Dutch
amaryllis is an example of a plant that goes dormant in response to dry
conditions.
A rhizome such as German iris
is essentially the main axis or stem of the plant in a horizontal position
under ground. Its development is less
complex than that of bulbs and corms.
Leaves and bloom stalks arise either from the growing apex or from nodes
farther back. Following bloom, that
particular section of the structure disintegrates, but others have been
developing to take its place. The differences
between rhizomes and stolons are not clear. Some
authorities consider any cylindrical horizontal stem a rhizome if it is normally
found below the soil surface, or a stolon if it is
above ground. However, in describing the development of tubers as in the case
of Irish potatoes, the tuber is most often considered the swollen subapical portion of a stolon. We take advantage of the food storage
function of potato tubers by including them in our diet. In order to propagate potatoes, we cut up the
tubers to include at least one "eye", which is actually a node
containing a bud. Each piece should
contain enough stored food to supply the new shoot until it can function on
its own. New roots arise adventitiously
from the base of the new shoot. What we
have done is propagate by taking stem cuttings. The original tuber is often
referred to as a "seed" potato.
Obviously, it is not a seed, which is produced in the ovary of the
potato flower.
Although these specialized
stem structures can be divided so the pieces contain buds that will develop
into new shoots, storage roots, on the other hand, do not have nodes and
buds. Sometimes adventitious shoots will
develop if the roots are subjected to proper conditions. Sweet potatoes (true roots) are subjected to
high temperatures, which seem to overcome what is called "proximal dominance,"
and shoots are then produced along the entire length of the tuberous
roots. However, in many cases no shoots
are produced unless a portion of the crown, the area where stem and root tissue
meet, is included with the root. This
propagation method is referred to as crown division. Dahlia is an example of a plant that must be
propagated in this way. Unless the
tuberous root is cut so that the top end includes a portion of the crown with a
bud, no shoots will develop. Stored food
in the root is used in the development of the new shoots. During the growing season, new storage roots
are formed at the base of the new shoot as the old ones gradually
disintegrate. What we have, then, is a
biennial root (living through two growing seasons) and an annual top (living
through one growing season). It should
be noted that fibrous plant roots behave somewhat differently and often can be
used for root cuttings, which will develop shoots. Shoot production occurs readily in many
species. It is easy to root plants that
"sucker", that is, normally send up shoots from the roots.
Wide variations of the basic
patterns of development are found in the different plant species. It is important to learn as much as possible
about each plant with which you work.
Frequently, questions arise about the possibility of forcing spring
flowering bulbs into bloom in the house.
Temperature control is very difficult in the average home, and results
are rarely successful. Most florists buy
preconditioned bulbs prepared by specialists using precise temperature regimes
required for each species. Only then
can bloom be expected to compare with that found when the plant is growing in
its native habitat.
Outdoor planting of crocus
and narcissus, including the large trumpet daffodils, can be done from
September until the ground freezes.
Results with these are quite satisfactory in most of our area except for
the extreme southern regions.
Tulips and hyacinths are
somewhat less successful and are best treated as annuals. Scales on the lily bulb are less protected
than are those of tulip, hyacinth and narcissus and must be handled more
carefully. They should not be allowed to
become dry. They are very subject to
fungus and virus diseases, so are very difficult to maintain successfully in
the warm, humid Southeast. Iris, daylilies
and anemones are good garden flowers for the Southeast. All underground storage structures are
subject to fungus and other soil-borne diseases. It is very important to plant in well
drained, sunny locations. In heavy clay
soil, do not plant as deep as is recommended for soils containing more sand or
organic material. A large dollar volume
of plant sales in the fall comes from sale of spring-flowering bulbs. It is helpful and important to know something
of their growth habits and requirements.
PLANTS WITH SPECIALIZED STEMS OR ROOTS
I.
Spring flowering. Hardy, planted
in the fall.
A. Amaryllidaceae - Amaryllis family
Bulbs
Allium sp.*
- Ornamental onion
Galanthus nivalis - Snowdrop
Leucojum vernum - Spring snowflake
Narcissus pseudo-narcissus - Daffodil
N. incomparabilis
- Short cup narcissus
N. jonquil - Jonquil - usually
fragrant
N. tazetta -
Polyanthus narcissus
N. poeticus
- Poet's narcissus
B. Iridaceae - Iris family
Bulbs
Iris reticulata
- Early fragrant iris
Iris xiphium
- Spanish and Dutch iris
Corms
Crocus vernus
- Large Dutch crocus
Crocus - Many others, beginning in
January
Rhizomes
Iris germanica
- German, or tall-bearded, iris
C. Liliaceae - Lily family
Bulbs
Hyacinth orientalis
- Hyacinth
Muscari spp. - Grape hyacinth
Tulipa gesneriana - Common garden tulip
Tulipa fosteriana and hybrids - Include large 'Red Emperor'
variety
Tulipa kaufmaniana - Water-lily tulip
Tulipa
species - More than 150 others
Rhizomes
Convallaria majalis - Lily-of-the-valley
D.
Ranunculaceae - Buttercup family
Tuberous root
Anemone coronaria
- Poppy anemone
A.
Blanda - Sapphire anemone
II. Summer and fall flowering (1. Hardy.
Fall planted).
A. Amaryllidaceae - Amaryllis family
Bulbs
Lycoris squamigera - Pink surprise lily
L. radiata -
Red surprise lily
B. Iridaceae - Iris family
Corms
Crocus autumnale
- Autumn crocus
C. specious and others - Fall
flowering crocus
C. Liliaceae - Lily family
Bulbs
Lilium
regale - Regal lily
L. candidum
- Madonna lily
L. speciosum
- Includes variety 'Rubrum'
Many more Lilium
species and hybrids, often difficult to grow in
the Southeast.
Corms
Colchicum species
Rhizomes
Liriope sp.
- Turf lily
Tuberous root
Hemerocallis
fulva - Day lily
III. Summer and fall flowering (2. Tender.
Dug and stored in fall, planted
spring).
A. Amaryllidaceae - Amaryllis family
Bulbs
Amaryllis vittatum
- Large flowering amaryllis
Hymenocallis
B. Araceae - Arum family
Tubers
Caladium bicolor - Elephant ear and
caladium
C. Cannaceae - Canna family
Rhizomes
Canna generalis - Canna
D. Compositae - Daisy family
Tuberous root
Dahlia pinnata
- Dahlia
E. Iridaceae - Iris family
Corms
Gladiolus gandavensis
- Gladiolus
*sp. - species, meaning many
different ones; some may be of unknown parentage.
LEAVES
The most conspicuous organs
of plants are their leaves. Leaves are
varied as to their arrangement on the stem, their form, the distribution of
their veins (venation), their structure, and many other characteristics. Leaf characters are used extensively in the
description of the species; however, reliance cannot be placed on them alone,
since their appearance is easily changed by environmental conditions. Generally, leaves of dicotyledonous plants
vary from leaves of monocotyledonous plants.
A typical leaf of a dicot* generally has
netted venation, one or more prominent veins with branches forming a
conspicuous net, while monocots* have parallel veined leaves with inconspicuous
branching between the veins such as in corn.
(*Accepted abbreviation for dicotyledons and monocotyledons plants,
respectively.)
Leaves have two main functions,
photosynthesis and transpiration.
The leaves on most plants are
the chief food manufacturing organs through the process called
photosynthesis. Photosynthesis may be
briefly defined, as the process whereby carbon dioxide (CO2) and
water (H2O) are transformed by energy from visible light absorbed by
chlorophyll (green pigments). During
daylight hours, this process of food manufacture (photosynthesis) results in
oxygen being given off by the plants.
The clean, fresh air in parks and planted areas is what makes them so
pleasant. We are dependent upon green
plants for the oxygen we breathe.
It is through the leaf that
almost all of the water is given off to the atmosphere by the aerial parts of
the plant in the process of transpiration.
The water balance of a plant is determined by the relative rates of
absorption through the roots and transpiration from the leaves. If loss of water is more rapid than absorption,
a water deficit occurs. A deficit in any
part of a plant may cause wilting, mostly seen in leaves.
The conducting tissue, xylem
and phloem, makes a continuous pipeline from the roots through the stems and into
the leaves. The veins, which form a
network throughout the leaves, are part of this pipeline. Water movement from the roots through the
vascular system to the leaves depends on lower water pressure at the top of the
plant. This pressure difference occurs
as a result of transpiration (loss of water from leaves).
In addition to providing the
mechanism for water movement through a plant, transpiration serves another
purpose. During the process, evaporation
of water from the leaf reduces leaf temperature and helps prevent injury from
excessive heat build-up.
FLOWERS AND CONES
Horticulturally important plants, with few exceptions, produce seeds.
This characteristic puts them in the most highly developed group of the plant
kingdom. This group is subdivided into
two groups: the angiosperms meaning
"covered seed," and the gymnosperms, meaning "naked
seed." Plants in the first group
bloom and develop fruit which encloses the seed, while those in the second
group have no true flowers or fruit but bear their seeds uncovered in cones of
similar structures. This group includes
the conifers, such as pines, the ginkgo, and a few others of importance to us.
Conspicuous because of their
bright colors, flowers come in many sizes and shapes as well as colors. Their principal value to the retail
nurseryman and the customer is ornamental.
Flowers are also used extensively in the description of the species. Their structure is less visibly affected by
environment than is the structure of leaves and stems.
The flower may be thought of
as a specialized stem with leaves adapted for reproductive functions. The following structures are present in a
flower that has all the characteristic parts:
1.
An outer set of green floral leaves (sepals), which enclose the other
parts of the flower until these are nearly mature. Collectively, the sepals comprise the calyx.
2.
An inner set of colored or white floral leaves (petals) constitute the
corolla. In many flowers, the petals are
showy and may aid in attracting the attention of insects that assist in
pollination.
3.
Within the petals, one or more sets of stamens (staminate or
"male" flowers). Stamens are
composed of pollen-bearing anthers supported by a filament. When the pollen is mature, it is discharged
through the ruptured anther wall and is carried by insects or the wind.
4.
At the center of the flower, one or more pistils (pistillate
or "female" flowers). Pistils
consist of an ovule-bearing base (or ovary) supporting an elongated region (or
style) whose expanded tip (or surface) is called the stigma. The ovule gives rise to the seed. The mature ovary becomes the fruit.
5.
These flower parts are borne on an enlarged portion of the
flower-supporting stem called the receptacle.
Pollination takes place when the pollen grains are transferred from the
anther to a stigma. Some plants are
self-fertile (Italian plum), and are able to use their own pollen, or pollen
from another plant of the same type.
Others (many cherries and
apples) need pollen from an entirely
different variety (cross pollination).
In addition, some species have male and female flowers on separate
plants; and in order to produce fruit, the female plant must have a male plant,
or pollinizer, nearby -- hollies and aucuba, for example.
A cone can also be considered
a specialized stem with leaves adapted for reproductive purposes. However, it does not have the typical flower
parts. Ovules and pollen are borne on separate cones, which are quite different
in appearance. The conspicuous cones are
the ovule bearing, or female cones. The ovules lie on one of the modified
leaves, called a cone scale, and are not surrounded by ovary tissue. Pollen is borne on much smaller cones and is
carried by wind to the ovule. Following
pollination, the scales of the cone close so that the ovule is protected while
the pollen grain germinates and fertilization of the egg cell occurs. This may take as long as a year. When the ovule matures into a seed, the scales
again open and the seed is released as from a pine cone.
FRUITS
The botanical term
"fruit" refers to the mature ovary and other flower parts associated
with it. It may include the receptacle
as well as withered remnants of the petals, sepals, stamens, and stylar portions of the pistil. It would also include any
seeds contained in the ovary.
SEEDS
A seed is a miniature plant
in an arrested state of development.
Most seeds contain a "built-in" food supply. Structurally, the seed is a matured ovule, although
various parts of the ovary may be incorporated in the seed coat. Plants naturally perpetuate themselves from
the seeds that develop from their flowers.
In the nursery industry, we are concerned with the flower itself and its
color, beauty and fragrance, rather than the seed or fruit produced by plants
such as azalea and marigold. We do like
to see fruit on pyracantha and holly. Many of our plants have been bred so that
they either produce unreliable seed or may not produce seed at all. This is especially true with annuals and
vegetables; the hybrid seed from which they grew is a result of controlled
pollinations. Were a person to collect
and plant the seed from a hybrid petunia or tomato, the resulting plants would
resemble the original parental types without many of the desirable
characteristics of the hybrid.
The fact that seeds are
really whole plants in a miniature form and in a dormant state will help you
understand why seed loses viability (ability to grow) and are no longer
good. The "miniature plant"
has actually "outgrown" its dormant period without receiving
conditions for continued growth (planting) and, therefore, has died. The period of time different kinds of plants
will stay alive in the dormant (seed) stage ranges from a few weeks to many
years. Package dating is used by most
seed firms to prevent sale of "old" seed. The proper care of the seed lengthens its
life considerably and we now see grass seed in vacuum-packaged containers,
which add months to its keeping quality.
Conversely, seed racks exposed to dampness, heat and temperature
extremes usually reduce the time seeds on the rack will remain viable.
The life-cycle of a
seed-bearing plant is divided into two broad phases, vegetative and
reproductive. The vegetative phase consists of two stages, the germination of
the seed (or rooting of a cutting) and vegetative growth. Although the germination phase may take only
a few weeks, the period of vegetative growth may last for years, with repeated
growth cycles.
The reproductive phase begins
with certain physiological changes indicating flower, or cone, bud induction,
followed by flower bud initiation and development, then flowering with the
subsequent production of fruit and seed.
VEGETATIVE REPRODUCTION
Many
plants reproduce by means other than seeds such as runners, tuberous roots, tip
layering, and underground stems. Man
takes advantage of these methods and adds a few others such as budding and
grafting to produce either more of the same plant or a new combination of two
or more plants.
Layering -- The development of new plants
by roots forming on a stem that is still attached to the mother plant. The rooted part is then separated from the
mother plant. Strawberries and
raspberries are naturally occurring examples.
Layering is not generally a nursery practice.
Division -- Many plants increase in growth
at the crown and produce new shoots from the base. As pointed out earlier, these shoots form
roots and may be separated as single new plants, or the entire plant may be
separated into sections.
This
method is particularly valuable for the propagation of herbaceous
perennials. Division is usually done in
the fall or spring. Examples are chrysanthemum, shasta
daisy, and pampas grass.
Separation -- Bulbs, corms, tubers, and
rhizomes are all similar in being underground stems which can produce
additional plants. Tulips form small bulblets, lilies form scales, gladiolus form small corms,
and dahlias form new tubers. These may
be removed from the parent plant after blooming and then planted at the proper
time. It may take two or three years to
produce a mature plant from some of these immature parts.
Cuttings -- Most woody plants are
propagated by cuttings. This technique
or dinarily assures the grower of obtaining a new
plant just like the original in all respects.
Cuttings may be made from many parts of a plant, such as the stem, leaf
or root, and are taken from different plants at various times of the year. Most cuttings need a humid atmosphere, warm
soil temperatures, light and a moist, but well-drained soil medium. Plant hormones are often used to accelerate
rooting (Fig. 4). A mist bed is commonly
used to provide moist conditions.
Graftage -- This
is the joining together of two different plant parts in
that
they will fuse and continue growing as one unit. The upper portion is the scion, while the
under part is the stock (rootstock, understock).
Grafting is done to produce plants that don't reproduce well or
"true" by other means (selected dogwood cultivars, pecan). Special effects also are possible, such as
"tree roses" and weeping cherries.
Some graftage is necessary because certain
soil pathogens make it difficult to grow some plants on their own roots. Fruit trees nowadays can be artificially
dwarfed by the use of special rootstocks which impart a dwarfing effect on the
scion. Whatever the purpose of grafting,
a successful take depends on how well the cambium layers of the scion and stock
match up. Often, the graft can be
detected by a slight swelling or a difference in the two barks. Once a graft union has healed, any suckers
arising from the root stock should be removed.
Budding -- This is a type of
grafting. A growth bud from one plant is
implanted under the bark of another of a related variety. "Four way apple trees" are produced
this way.
GROWTH CHARACTERISTICS
One
of the questions asked by many of the customers coming into the nursery is
"how large (or how tall) does it grow?" While most trees and shrubs have a maximum
under good growing conditions, this growth may not be reach for many, many
years A large number of the trees and
shrubs can be maintained at much lower height or size by controlling growth
with selective pruning. For example, Photinia
fraseri
(Red Top) may eventually reach twenty feet in height; yet, this plant is
used in many locations where it is held to a certain height. A large number of the trees and shrubs can
be maintained at a much lower height or size by controlling growth with
selective pruning. For example (Red Top) may eventually reach twenty feet
in height; yet, this plant is used in many locations where it is held to a
height of 6 to 8 feet for a period of years.
Possibly, the best answer for the customer is to describe a plant as low
growing, medium or tall, and explain some of its possible variations. It should also be remembered that each plant
is an individual, and its response to environmental conditions may be quite
different from another plant of the same species growing nearby.
Many
of the plants found in our gardens and in the nursery originate in other states
and in foreign countries that have similar climate to our climate in the
Southeast. The more nearly one's garden
duplicates the climate of the plant's original home, the better the plant will
grow. The average gardener's great
problem is trying to grow plants that are not adapted to the conditions found
in his garden.
Each
plant found in nature has, over the centuries, adapted itself to certain
climatic conditions -- sunlight, temperature (day, night and seasonal),
moisture (rainfall and humidity) and soils (type, pH and nutrient
content). These factors combine to make
what is called the "optimum growing conditions." Some plants have a wide tolerance in all or
some of the requirements, enabling them to be grown over a wide area. Others are so demanding in their requirements
that they are greatly limited in usage.
Customers newly interested in gardening should be encouraged to start
with some of the more adaptable plants.
The
great challenge in the nursery industry is to provide as wide a variety of
plants as possible that can develop to their fullest potential in the
situations where they will be used. The
answer to the challenge is partly in understanding the plant's needs and
learning to adapt existing conditions to meet these needs.
Irrigation
can augment natural rainfall, soil amendments and fertilizer can modify the
natural soil and both shade cloth or trees can reduce light intensity. However, increasing light intensity is more
difficult. Selection of the
brightest areas is necessary for plants requiring high light
intensity (bright sunshine).
In
our gardens we will find certain consistent variations in
light, temperature and moisture. We
call these different small areas "micro- climates" (little
climates). For example, on the north
side of the house, there is more shade, the ground stays cooler and there
is less light intensity. The south
side of the house and the west side of the house have less shade (barring
trees), the ground warms up faster and to a higher temperature and the
light intensity is higher. The east side
is generally cooler than the south or west, yet warmer than the
north. Trees, structures and
prevailing wind patterns can modify these conditions somewhat. Elevation may also be a factor as cold
air drains to the lower areas at night, possibly even affecting the
survival of some plants that are not sufficiently cold tolerant. Micro-climates differ from the climatic
conditions of the general zone by only a few degrees in temperature, but this
may be enough to cause a plant to do well, to merely exist, or to die
completely.
PRUNING
Plant growth can be both
controlled and modified by proper pruning.
We have already mentioned the possibility of keeping a plant to a
desired height by pruning. The most
conspicuous example of this is found in formal hedges that are kept to size and
shape by regular clipping. It is
important to shape a formal hedge so that the top is narrower than the
bottom. If this is not done, the lower
leaves drop due to inadequate light, and the plant is bare and"leggy". Plants can also be kept to a size, yet remain
informal in growth pattern. Abelia grandiflora generally grows as a large shrub, which may
get too big for many residential sites.
It can be maintained as a much lower informal shrub by selectively
pruning out leaders, beginning when the shrub is about three and
one-half feet high. These leaders are
cut back to below the height of the surrounding foliage, leaving an
attractive plant without visible stubs.
Pruning can be used to
produce a fuller shrub or tree by encouraging branching. Pruning can increase the light entering into
the center of the plant by thinning; it can aid in disease prevention by
allowing more air passage within; it can reduce or increase flowering or
fruiting; shape the plant formally or informally; rebalance growth after
a poor start or after injury and help the plant through periods of stress. In short, pruning is one of the best
tools we have to aid plants to reach the optimum growth to fulfill our desires
for it. Even if we don't prune, nature
will do some of it for us, though not necessarily in a desirable fashion.
Gardening is the science, art
and fun of learning the various kinds of plants and their growth requirements,
then adapting conditions to meet these plant needs. The nurseryman needs an ever-increasing
amount of this knowledge to best serve his customers. Take some time each day to observe the
different plants in the nursery, get to know them and their characteristics
throughout the year and read as much about them as possible.
For further information,
consult any book on general horticulture, plant science or botany.
SUMMARY
Competence in technical
knowledge of plant growth aids the nurseryman in growing plants and answering
customers' questions in language that they most easily understand.
In general, all plants are
made up of many parts:
-- roots (root system), which anchor the
plants, absorb and conduct water and nutrients, and
may store food.
-- stems, which support the plant parts,
conduct water, nutrients and plant food and may store
food.
-- leaves, which are the site of
photosynthesis and transpiration and are also ornamental.
-- flowers (or cones), which may be
ornamental and produce fruit and seeds used in
reproduction.
Plants are very diverse in nature,
yet are grouped together by function and characteristics that they have in
common.
Knowledge of plants aides us
in choosing the right plants for the right location and in giving the proper
care to the established plants.
WATER AND
WATERING
Water is one of the most
important factors in regulating plant growth.
Water is also one of the most misused commodities. Cases of underwatering,
overwatering, wrong method of watering, wrong time
of watering and using the wrong type of water easily exceed those of proper
watering. The average individual
probably does not realize the number of mistakes that he makes because of
the many factors affecting water and watering of plants.
In order to better understand
just what is involved in proper watering, we need to know something about
the nature of water and how its use is affected by the plants being
watered, the soil (or media), methods of application, and in some cases,
the nature of the containers used.
Sources of water include rain, private wells, municipal or
commercial water systems (using well water, lake, reservoir or river sources). Cost of water usually depends upon the
source and varies considerably from area to area. However, this economic factor is important
enough, along with the ever-increasing demand for available water to make
use of proper watering procedures practically mandatory.
With water coming from so
many possible sources, its quality varies considerably. For example, the soluble salt content may
vary from practically none (rainwater) to high soluble salt content. In some areas where the soluble salt content
is not too high, there may exist problems of a single nutrient being too abundant. More often when total soluble salts are
high, those nutrient elements that are already present should not be
applied as fertilizer. Where an
element is already at toxic quantities in the water, avoid using more of
that element in the fertilizer applied and choose plants more tolerant to
the salt. Acidic water seldom causes any
serious problems, but high pH (alkaline) may be a real problem. Where the soil pH is already over 7.0,
the addition of alkaline water could well be detrimental to
plant life. Watering through a
mulch of Canadian peat moss should help alleviate the problem.
In some areas of hard water
(either alkaline or from some other chemicals), the homeowner often has a
water softener. Water from this type of
water softener should never be used to water plants, either indoors or
out. These softeners use sodium ions (NA+) to replace the ions that cause
the hardness. Sodium is not a plant
nutrient but may enter the plant instead of potassium (essential to plant
growth) causing a potassium deficiency in the plant. Where softening of the water is the only
alternative, the customer should follow the example of the commercial
plant growers and use a system that uses phosphoric acid as a softener.
WATER APPLICATION TO LAWNS AND PLANTS IN THE GROUND
When water is applied by any
means, rain, sprinkler or flood irrigation, only a fraction is actually
used by the plants. The first water lost
is due to run-off from the soil surface.
Run-off increases the surface crusting, compaction of the soil as
well as increased slope of the land.
Soil compaction and surface crusting can be relieved by the
addition of organic matter to the soil and even by surface mulching in
extreme cases. If the slope of the
land is too great, water must be applied slowly, or in the case of rain,
adequate drainage provided. Water
penetrates the soil and fills up all of the pore spaces (spaces between
the soil particles). The rate
of penetration depends upon the type of soil, with penetration in sandy
soil much more rapid than in clay soils.
For example, when one inch of water is applied (excluding run-off),
sandy soil will be penetrated 12 inches; loam, 6 to 10 inches; and clay, 4
to 6 inches.
Some of the water that
penetrates the soil continues to move downward due to the force of
gravity. Soil with larger particles
(i.e., sandy) has larger pores and, therefore, loses a larger amount of
water to gravity. While this water
is theoretically available to the plant, it remains in the soil for such a
short time that it is of negligible importance.
Nutrients are also carried down past the plant roots in this water
movement. When all of
this gravitational water has been removed, the soil is said to be at
field capacity. The percentage of
water remaining in the soil at field capacity, based on dry weight of the
soil, ranges from 7 percent for sand, through 15-25 percent for loam, to
35 percent for clay soils. However, all
of this water isn't available to the plant as percentages varying from 3.5 percent
for sand to 17.5 percent for clay are held so tightly to the soil
particles as to be unavailable to the plant. Organic matter added to sand both increases
the percentage of water at field capacity and the amount of bound
water. The net effect is to increase
the amount of water available to the plant.
There are two forces that
combine to hold the water in the soil, adhesion and cohesion. Water adheres to the surfaces of the soil
particles. Water also has a high
surface tension causing the water droplets to cohere (hold together). This can be partially demonstrated by
spilling a small quantity of water on a clean pane of glass. Some of the smaller droplets tend to
join together to form larger droplets.
This is cohesion. If the pane of
glass is tilted slightly, a film of water will remain as the rest runs
off. This film is an example of
adhesion. A further demonstration that
also shows the effect of pore size is to take several glass tubes of
different diameters and place one end in a basin of water. The water begins to move upward in the
tubes, moving highest in the tube with the smallest diameter.
WATERING PLANTS IN CONTAINERS
Containers vary from those
for the smallest house plants to some extremely large outdoor containers
designed for trees or large shrubs.
However, they can be classified for watering purposes into two
groups; those with drainage and those without it. It is essential in either case that the
plants in the containers be planted in a proper media. There are very few instances where
a native soil is satisfactory.
Normally, organic matter must be added.
The best situation for the container plant is to prepare a media
using sand and organic matter (usually peat or ground bark) or loam soil,
sand and organic matter. The medium
may vary according to the plants, but should contain from one-third to
one-half organic matter by volume (possibly even more for azaleas,
camellias and some house plants). This
media is not only better for the growth and support of the plants, but greatly
reduces watering problems.
Water should be added slowly
to the container to avoid splashing and washing out of materials from the
surface, the rate somewhat regulated by the size of the container. Where the container has natural drainage,
apply sufficient water so that a small amount comes through the drainage
holes. Care should be taken that the soil ball never dries out or it will
separate from the surface of the container and the water will simply flow
out around the soil ball and through the drainage holes, leaving the
interior of the ball dry. Where the
container does not have drainage, a measured amount of water (determined
by practice) should be added to the container.
This can be done by timing the flow of water or using a watering
can with a measured volume. Further protection to prevent "wet
feet" of the plants in containers without drainage is provided by
placing a small amount of activated charcoal scattered on the bottom.
The most difficult question
of all is "when to water (how often?)". The only proper answer is "when the
plant is dry." However, extreme
care must be taken in answering this question for a customer to not sound
facetious. A good indication of
need for watering is the surface drying of the soil from a depth of 1/4
inch to one inch, depending upon the type of plants and root depth. Another test is to form a hand-full of the
soil in question into a ball in the hand.
If it adheres together, there is usually adequate moisture; if it
appears more than a "mud pie" there is far too much moisture; and if
it completely crumbles, there is insufficient moisture. By determining where the ball fits in
this scale, the time or interval of watering can be determined. With the
exception of those plants that grow in water or on the very edge of ponds
and lakes, it is generally better that some drying takes place
between watering, short of wilting.
So, water less often and water enough to penetrate the soil to a
depth of the roots. This may mean daily
(or more often in extreme dry weather) for plants in small containers, at
4 to 8 day intervals for lawns and shallow rooted plants (depending on
soil type), and up to monthly intervals for trees and large shrubs.
This method of watering is
much less important, providing the amount of water is right and that
there is no surface splashing or too great a run-off. However, several
methods are available, some of which are time saving and possibly water
saving. Grandmother may have gotten
satisfactory results from sprinkling her lawn every night with a hose
nozzle; but then grandmother was generally there every night to sprinkle
it. Her grass has extremely
short roots (roots only grow well to the depth of water penetration) and
had she neglected it for several days, it would have suffered badly and
died within a week or two. Hand
sprinkling one inch or more of water on the lawn at one time at the proper
rate would exhaust the patience of most of us.
Hand watering is satisfactory for plants in containers and
occasionally supplying extra water to a tree or shrub in or near a lawn
area where normal lawn watering does not penetrate deep enough. Even hand watering can be speeded up by
using water breakers or bubblers on the end of the hose. These allow greater volume of water at
reduced force.
Much of the watering of individual gardens
is done by using movable sprinkler-hose combinations. With several sizes of hoses and many
different sprinklers available, it is imperative that the right
combination of hose and sprinkler placement be used. Hose sizes generally available are 1/2
inch, 5/8 inch and 3/4 inch, with 7/16 inch and 1 inch occasionally on the
market. The 7/16 inch is not adequate for most situations and 1 inch is
too heavy for most homeowners. The
most satisfactory sizes are 5/8 inch and 3/4 inch, with the 3/4 inch
adequate for watering containers (except the larger sizes) or for small
sprinklers to catch odd corners. A
customer should be advised to purchase a quality hose for greatest
satisfaction. Cheaper hoses tend to be stiff
and difficult to manage in cold weather, tend to kink, crack, split
near the ends, and burst when under pressure too long in the hot sun. The most common types of sprinklers are
the rotating impulse sprinklers (rainbird type),
oscillating and fixed head sprinklers.
All sprinklers have particular spray patterns, most of which require
overlapping to provide the proper amount of water evenly distributed over
the entire area. A good method of
checking evenness of watering is to secure a number of straight sided cans
of equal diameter (coffee cans are good), space them evenly throughout the
area and water the whole area using your normal placement. The result should be the proper depth of
water in the can with the shallowest level and with no more than a slight
deviation between the can with the shallowest level and that with the
greatest amount of water. If any area is
receiving inadequate water or if the distribution is too uneven, the
pattern of sprinkler placement and/or type should be changed.
Permanent sprinkler systems,
with either manually operated valves or under automatic controls, take
much of the guesswork out of sprinkler setting when properly
designed. They also save labor and
eliminate hose marks in the lawn as well as unsightly coils of hoses lying
around. They are available in black
plastic, PVC and galvanized pipe, with either impulse type, rotating
or fixed head sprinklers. However,
except for the simpler installations, and unless the customer is
experienced, they should be designed and generally installed by
experts. If your nursery provides a
sprinkler designing and/or installation service, direct your customer to
that department. If not, determine
your nursery's policy of referral. Do
not attempt to design a system for the customer yourself if you are not experienced
in this field, nor encourage your customer to proceed without adequate
assistance unless he has had experience.
For those systems the customer can install himself, the coffee can
method of checking results in a good policy to insure
proper coverage.
For further information on
watering, the reader is referred to:
Sunset Basic Gardening Illustrated, pages 45-50; Sunset Western
Garden Book, pages 44-48; reference manuals produced by Irrigation
Technical Services of Lafayette, California.
SUMMARY
Water is one of the most
important factors in plant growth. Yet
water is probably one of the most misused commodities and proper watering
is one of the most difficult concepts to convey to the consumers.
There are many more cases of
improper watering (too little, too much, wrong methods, wrong timing,
etc.) than of watering properly.
All plants need a certain
quantity of water for proper growth functions. The amount of water applied
to supplement natural rainfall, and the frequency of application, must
take into account the factors of plant growth, transpiration, run-off, and
the water-holding capacity of the soil.
Plants in containers, especially those containers without a natural
drainage, need additional special care in watering.
Methods of watering vary
greatly, from hand watering with hose or watering can, use of portable
sprinklers and garden hose, to permanent sprinkler systems. Adequate coverage and proper timing are more
important than the method used.
While permanent systems
generally are best, they should be designed by experts.
SOILS
What is soil? Good soil is a living, breathing, complex
moisture of minerals, organic matter (dead organisms) and living organisms
(as many as two billion per cubic inch).
It provides support for plant roots and holds moisture and minerals
for plant use. It has been formed over
the centuries by the actions of water, wind, sun (heat), freezing and
thawing and by the action of plant life itself. It varies in thickness from a few inches to
many feet. It also varies in texture, structure and mineral composition
based on its parent material and the conditions under which it was
formed. Rarely is any one soil
completely satisfactory for all of the plants available in nurseries.
Soil texture: The relative size of the particles that make
up the soil is expressed by the term texture. It refers to the fineness or coarseness
of the soil. More specifically,
texture is the relative proportions of the different soil separates, sand,
silt, and clay. Many physical and
chemical reactions in soil are governed by its texture. This is because texture determines the
amount of surface on which the reactions can occur. Soil particles have been divided into
groups entirely on the basis of size, without regard to their chemical
compositions, color, weight, or other properties. These sizes range from
slightly less than 1/8 inch diameter for fine gravel down to clay
particles of less than 1/12500 inches.
Size and Surface Characteristics of Soil Separates
To clarify a relative
particle size, imagine that the clay particle is the size of a radish
seed. In comparison, a single silt
particle would be the size of a basketball and a coarse sand particle the
size of a 7-1/2 foot (diameter) balloon.
The presence of the three
separates can be determined in a soil sample by rubbing a small amount of
moistened soil between the finger tips.
Sand, being granular, has a gritty feeling, silt a slick feeling (like
talcum powder) and clay a slippery-sticky feeling.
Soils are named on the basis
of the separates they contain; i.e., "sandy soils", "clay
soils", "clay loams", "silt loams", etc. with the
name indicating the predominate particles present. A proportion so that none of the three
was dominant over the other two is a loam.
The term loam is also much misused (i.e., garden loam), so that it
is essential to have a firmer basis in discussing soil problems with a
customer. It is best where
any doubts exist to have the customer bring in a sample of the soil from
his lawn or garden.
Soil Structure: The way that the various particles are
arranged within the soil determines the soil structure. Structural names, i.e., sandy,
platy, crumby, blocky, are self descriptive. Structure is either single grained as in
sand soils or in aggregates of smaller particles grouped together as in
the clay soils. Soil structure
affects water movement and the handling qualities of soils.
Organic Matter: Soils are classified as either mineral soils
or organic soils, according to the percentage of organic matter present in
the top 12 inches. Organic soils
are those which have more than 30 percent organic matter (20 percent if
the soil is loamy sand or coarser).
Organic soils are further divided into peats
and mucks, depending on the degree of decomposition. In peat, most of the original plant fibers
can be recognized; whereas, in mucks, they are decomposed beyond
recognition. Most soils
have considerably less than 20 percent organic matter and, in fact, most
soils need added organic matter for best plant growth.
Organic matter favorably
affects soil structure, increasing the water holding capacity of sandy
soils and the workability of the clay and other "heavy"
soils. (The terms heavy and light
soils, according to common usage, are based on the effort needed to move
tillage equipment through the soil, not on the physical weight of the soil
itself).
The organic matter in the
soil is the residue from plant and animal tissue. The presence of living organisms in the soil
is also important, contributing to the better growth of plants. Some of these organisms cause the decay
of dead plant and animal material, releasing nutrients which
become available to living plants.
Others fix nitrogen from the air (nitrogen fixing bacteria), attack
harmful organisms, burrow tunnels through, improving aeration and
releasing minerals (earthworms), and change inorganic (mineral)
soil constituents.
Some of these organisms in
the soil are also detrimental to plant life. Fungi cause rots, damping off
and other diseases; bacteria cause blights; cutworms eat plants; nematodes
damage roots and other plant structures; and some forms of insect life in
the soil are also detrimental. Soil in
good tilth and properly cared for often contains
beneficial organisms that hold the detrimental organisms in check. However, when the number of
detrimental organisms of any type build up to the point where natural controls
cannot cope with them, chemicals must be used to bring them back into
balance.
Soil Reaction: Refers to the acidity or alkalinity of the
soil. It is expressed in terms of
pH, in units on a scale of 0 to 14, with 7.0 neutral, below 7.0 acid (increasingly
acid to 0). For example, a pH of 5.0 is
ten times as acid as a pH of 6.0; and above 7.0 alkaline in reaction
(alkalinity increases to 14 the same manner as acidity increases below
7.0). The greatest number of plants
grown in the Northwest do best in a pH range of 5.5 to 7.0 with some
plants such as azaleas, rhododendrons and camellias preferring
soil slightly more acid (pH 5.0-6.0).
The soil pH is not the factor
which controls plant growth. It is only
a good indicator of what nutrients are soluble (available) or insoluble (unavailable),
which type of micro-organisms are likely to predominate and what type of
chemical reactions to expect. It is a
useful tool when backed by knowledge and experience. Simple test kits for soil reaction (pH) are
readily available and are sufficiently accurate when used carefully. They may well be a useful tool to
the nurseryman to help solve a
customer's soil problem, so long as other pertinent information about the
soil is also available.
Soil reaction can be modified
by addition of certain soil additives and by use of fertilizers. The addition of lime will make acid soils
more alkaline (also wood ashes).
The addition of gypsum or sulfur will make alkaline soils more
acid. Care must be taken to prevent an
over-reaction. It is often
advisable to base corrective measures on a competent soil test unless previous
experience with the soil type in question is adequate.
Soils contain varying amounts
of natural fertility based largely on the parent material from which they
were formed, their texture, and the effects of moisture on them. Clay soils usually contain more minerals than
sandy soils. Clay soils have a much greater surface area than sands. Minerals are usually held on the surface
until released to the plants on contact with the plant roots. The addition of organic matter to a sandy
soil increases the surface area and improves the soil's ability to hold
minerals. Even though clay soils contain
more minerals than sandy soils, it cannot be assumed that any or all essential
nutrients are present or absent in any given soil sample. Only soil testing by a reputable
laboratory can indicate this. What we
can assume is that in adding a chemical fertilizer, it will be immediately
and totally available in sandy soils, but will also leach out rapidly in successive
waterings. In
clay soils (or sandy soils modified with sufficient organic matter) much
of the nutrients will adhere to the surface of the soil particles to be
more slowly available to the plant and less subject to leaching or loss.
The customer's soil
problems: As we have already noted,
soils vary greatly. Even next door
neighbors in the same valley area could have different types of soil, as much
of the soil adjacent to rivers had been deposited at various times and in
various thicknesses. It is difficult
to determine the nature of a customer's soil from a color or word
description. Black soil could equally describe a loam (an excellent soil
with almost no problems) to one of the clay soils found in some areas. If possible, suggest that the customer
bring in a sample of his soil when you have any doubts in your mind about
making recommendations for plantings, additions, etc. At least, ask enough pertinent questions
to assure that you understand his problems.
According to the size of the
area in question and to the specific plants that the customer is
interested in, we can consider soil treatments and usage in three
ways. The first consideration is those
small areas that require special treatment; i.e., planter strips, built-in
planter boxes, containers and sections of larger areas that are to be used
for plants like azaleas and camellias that have more demanding soil
requirements. The best suggestion for these
situations is the complete removal of the soil (to a depth of at least 12
inches) and replacement with a soil mix suitable for the intended
plants. For very small areas and planters, planter mixes are usually sold
in the nursery. For larger quantities,
a suitable mix can be made based on the original soil plus additives. Mineral additives include sand,
vermiculite, perlite and pumice and become a
permanent part of the soil. Organic
additives include peat, ground bark, sawdust, leafmold,
manure, etc. These break down to an
end product called humus, and should be added on a regular basis. A good rule of thumb for a loam soil is
the addition of 1 part sand and 1 part peat or ground bark to 2 parts of
soil. Heavier soils would require
greater quantities of sand and bark, while sandier soils would only need
the organic matter additives.
The second situation would be
that of larger areas where the soil has no special problems but needs
improvement in water-holding ability and/or increased organic matter. Generally, organic additives are recommended
here, either over the whole area when lawn, ground cover or bedding plants
are to be used, or in enlarged holes for the planting of shrubs and trees
from containers. At least 1 inch of
ground bark or peat should be added to most "good garden soils"
up to a minimum of 25% (3" or organic additive mixed with 9" or
"plow depth" of soil) to those soils that are very sandy or have
too much compaction or other problems.
For example, a 5.5 cubic foot (compacted) soils that are very sandy or
have too much compaction or other problems. For example, a 5.5 cubic foot (compacted)
bale of peat moss equals 11 cu ft. of loose material. This will cover an area of 131 sq. ft. (10 x
13), one inch deep. One cubic yard of
ground bark will cover an area of 108 sq. ft. (approximately 10 x 11), three
inches deep.
These additives must be
thoroughly mixed with the soil. The
greatest problem in planting canned stock is the transition layer between the
soil ball in the container and the native soil surrounding the place where it
is planted. If the shrub or tree is
planted into a hole barely large enough to hold the soil ball, there is a good
chance of problems occurring. If the two
soils have too great a difference, water may not be able to move from one to
another, either creating a situation where the soil ball becomes overly wet
while the surrounding soil remains relatively dry, or the converse, where the
surrounding soil remains moist and the soil ball drys
out. Either situation can cause damage
to roots and the failure of the plant to establish itself. If the native soil is either considerable
heavier or more compact than that in the root ball, the plant roots may not be
able to penetrate the native soil. All
of these problems can be resolved by establishing a proper transition layer
between the soil ball and the native soil.
The hole for the plant is dug at least twice as large and half again as
deep as the soil ball. The native soil
removed is mixed, 2 parts native soil to 1 part sand and 1 part peat or ground
bark. This mixture is placed in the
bottom of the hole, firmed down to the proper depth and the tree or shrub is
placed in the hole. The same mixture is
used to fill the hole being sure to properly tamp and water it in.
The most commonly used organic additives are peat moss, ground bark, compost and miscellaneous organic materials. Pea